2.3.1 Geological mapping of coalfields Coalfields can be divided into two categories: exposed coalfields, where the coal-bearing strata outcrop at the surface, and concealed coalfields, where they are hidden beneath younger rocks. Exposed coalfields can be defined with considerable precision by surface geological investigations; indeed geologists recording field data still represent the cheapest exploration ‘tool’ available to the coal industry. In populated regions, the locations of coal outcrops are well
2.3 Exploring for coal Early miners would have found it easy to trace the distinctive black colour of coal along an outcrop (for example, a coastline or river valley), and surface trenches were used to locate less obvious outcrops. However, tracing an outcrop underground was problematical as the only means of exploration was by digging costly trial shafts. The development of exploratory steam-powered drilling in the early 19th century improved matters, but it was not until the mid- to late- 20th century that more a
2.2 Winning coal in former times Coal was probably first used as a fuel by early Chinese civilizations, and there is evidence for coal working in the UK since Roman times. However, early approaches to mining were limited by the available technology, and left much of the coal behind. At first, coal was dug from seams exposed at the surface in shallow excavations into valley sides that followed the coal seam. The amount of coal that could be extracted from these trenches and from adits (short horizontal tunnels) w
2.1 Finding and extracting coal Coal is often regarded as the principal fossil fuel, and with good reason. There is almost three times more energy available from the global proven coal reserves as there is from proven oil and gas reserves taken together. Therefore, it is unsurprising that even today much time and effort is spent locating it. This section considers the techniques used in coal exploration and how coal is produced from surface and underground mines. But first, a brief look at a few of the historical aspe
1.7.1 Carboniferous mires During the late Carboniferous, mires developed over vast areas of the UK. Much of today's land area was an extensive, low-lying plain bordering a sea to the south (a sea that was soon to be the site of a mountain-building episode). Any mountains that existed lay hundreds of kilometres to the north. Large river systems meandered southwards across these plains. At that time, the UK lay in tropical latitudes, almost on the Equator (see Author(s):
1.7 How old is coal? Not surprisingly, the distribution of coal deposits through time corresponds closely to the origin and distribution of land plants. (This is discussed further in Section 4.) Coals are commonly found in rocks from Carboniferous times onwards, Devonian coals are rare, and pre-Silurian true coals are never found. This coincides with evidence for the evolution of land plants, which first appeared in Silurian times about 400 Ma (million years) ago, colonized the land surface rapidly through the De
1.6 Impurities in coal Coal rank reflects the maturity of a coal, but another variable is the ratio of combustible organic matter to inorganic impurities found within the coal. As discussed earlier, impurities result mainly from clay minerals washed into the mire prior to its eventual burial. In addition, some impurities are formed from the plant material itself during coalification. These inorganic impurities are non-combustible and therefore leave an inert residue or ash after coal combustion. High-a
1.5 The physics and chemistry of coal formation Coal is a type of sediment made up mainly of lithified plant remains. But how does spongy, rotting plant debris become a hard seam of coal? As discussed earlier, plant material growing in mires dies, and then rots under anoxic conditions to form peat (by the process of humification). With time, the mire becomes covered with layers of sediment, the weight of which squeezes water and gas out of the pore spaces and compacts the vegetation. As subsidence allows deposition of further mireâ€
1.4 Coal-forming environments in the geological record Figure 5 simplifies a typical vertical succession of sedimentary rocks found in many coalfields. The sequence from the base of the section upwards reveals the following: When a mire starts to form, the first plants take root in underlying clays or sands that form the soil. Their r
1.3.2 Peat formation in raised mires Mires can also form inland within low-lying depressions, provided the rate of precipitation exceeds the rate of evaporation (Figure 4a). Peat is impermeable and so its accumulation progressively impedes drainage. This attribute gives mires the ability to maintain a water table independent of the area surrounding them.
1.3.1 Peat formation in deltas and coastal barrier systems Since mires require poor drainage, low-lying land close to coastal areas might provide the right conditions for peat to form. Most extensive areas of modern peat formation are indeed situated not far above sea-level, and as Figure 2 shows, they are commonly associated with river deltas and coastal barriers. Such enviro
1.3 Coal-forming environments today Coal formation begins with preservation of waterlogged plant remains to produce peat and then slow compression as the peat is buried. About 10 m of peat will compress down to form about 1 m of coal; clearly large amounts of plant debris must be available for preservation. Even so, for a significant thickness of peat to accumulate there must be a balance between the growth of plants and the decay of underlying dead material to form peat (a process known as humification). Su
1.2 The origins of coal If you examine a piece of coal, at first sight it appears black and rather homogenous. However, closer inspection generally shows a series of parallel bands up to a few millimetres thick. Most obvious are shiny bands that break into angular pieces if struck. Between them are layers of dull, relatively hard coal and thin weak layers of charcoal-like carbon. Coal splits easily along these weak layers, which crumble to give coal its characteristic dusty black coating. Microscopic examinati
1.1 Introduction There are many environmental reasons why coal is a rather undesirable source of energy. Burning it introduces large amounts of gases into the atmosphere that harm the environment in a variety of ways, as well as other, solid waste products. Coal extraction leads to spoil heaps and mines that scar the landscape, land subsidence that affects roads and buildings, and in some cases water pollution. With apparently so little going for it, why do we rely so much on coal to meet our energy nee
1 Natural groups Darwin made extensive observations on a great many creatures, including mammals, and noticed that species fell into natural groups, e.g. lions, tigers and leopards have many similarities, and resemble cats. On the basis of his observations, he was able to place mammals in distinct groups. His work has continued, and we now recognise that mammals have evolved from a common ancestor, and have branched into many different groups, or ‘Orders’. The animation below shows the different O
Learning outcomes By the end of this unit you should be able to: develop an appreciation of the huge variety of different mammals that exist on Earth today; see how fossil evidence can help us to understand evolutionary history; understand how the structure of DNA can help us to detect differences between different species; apply the techniques of DNA analysis to work out which mammals are most closely related to each other; appreciate t
Acknowledgements Grateful acknowledgement is made to the following: Except for third party materials and otherwise stated (see terms and conditions), this content is made available under a Creative Commons Attribution-NonCommercial-ShareAlike 2.0 Licence Figures 4 and 8 Alber
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7.1 Introduction In all eukaryotic cells, proteins that are destined for the plasma membrane or secretion are synthesised in the rough endoplasmic reticulum and enter the Golgi apparatus where they undergo a variety of post-translational modifications, before transfer to the cell surface in secretory vesicles. Which post-translational modifications of proteins occur in which compar 6.2 Endocytosis Fluid-phase uptake by pinocytosis can be broadly categorised according to the size of the endocytic vesicle and this also relates to how the vesicle is coated (Figure 35). The rate of internalisation is directly proportional to (i) the concentration of extracellular molecules, (ii) the volume enclosed by the vesicle and (iii) the ra 4.5 Summary Targeting sequences at the N-terminus of proteins direct translation across the ER, and act as signals for import to the nucleus, mitochondrion and chloroplasts. Sequences at the C-terminus control traffic through the ER and the Golgi and to peroxisomes. Glycosylation is directed by signal sequences that act as targets for N-linked glycosylation in the ER and O-linked glycosylation in the Golgi apparatus. Glycosylation and remodelling of polys