Samuel was the eldest son of William Mellish (1605-1667) of Stepney and Doncaster, and his wife Martha, née Goldsmith, and a grandson of Edward Mellish of London. He was baptised in the parish of St Dunstan, Stepney.
Samuel moved from London to become a barrister-at-law in Doncaster, Yorkshire.
In around 1670 he married Joan (d 1708/9, aged 75), daughter of Thomas Prowse and widow of Tobiah Harvey, citizen and vintner of London (who died before 1663). Joan had three children from her first marriage:
Samuel and Joan had:
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